The first report of lesbosoxylon süss & velitzelos from the early-middle miocene of eastern anatolia

Ünal Akkemik*, Dimitra Mantzouka, Demet Kiran Yildirim

*Corresponding author for this work

Research output: Contribution to journalArticlepeer-review

10 Citations (Scopus)


A new fossil pine species from eastern Turkey is described and its botanical affinities are discussed. The sample was collected from the city of Kemaliye, Erzincan province, Turkey, and derives from the early-middle Miocene Divriği formation. Transverse, tangential and radial sections were taken from the petrified wood, and its palaeoxylotomical features were investigated. Based on its anatomical features including idioblastic cells in rays a new fossil-species of the genus Lesbosoxylon Süss & Velitzelos was identified as Lesbosoxylon kemaliyensis Akkemik & Mantzouka, sp. nov. Diagnostic features of the new species are: Transition from earlywood to latewood mostly gradual; axial and radial resin canals with thin-walled epithelial cells present; latewood tracheids thin to thick walled; bordered pits on radial walls of tracheids 1-2(-3) seriate; crassulae common; rays heterocellular, uniseriate, partly biseriate; uniseriate rays up to 27 cells high; fusiform rays up to 30 cells high; axial parenchyma occasionally present; ray tracheids 2-3 rows; cell walls of ray tracheids smooth; cross-field pitting pinoid, 1-2(-6) pits per cross-field. Detailed investigation of the botanical affinities of the new fossil wood suggested that the most closely related modern species is Pinus canariensis C. Sm in Buch, a relict species from the Canary Islands.

Original languageEnglish
Pages (from-to)427-441
Number of pages15
Issue number23
Publication statusPublished - 2020

Bibliographical note

Publisher Copyright:
© Publications scientifiques du Muséum national d’Histoire naturelle, Paris.


has been provided to the second author (DM) by the Alexander von Humboldt Foundation.

FundersFunder number
Alexander von Humboldt-Stiftung


    • Biogeography
    • Conifer
    • Miocene
    • New species
    • Pinaceae
    • Sivas Basin
    • Wood anatomy


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